Addison returns to Seattle Grace to perform an operation and is surprised by all the changes in the lives of her former co-workers; Derek and Meredith's clinical trial has its first patient; Rebecca returns with shocking news for Alex.
Six weeks after Derek and Meredith ended their relationship, Meredith, Alex, Izzie and Cristina compete in a surgical contest, with Bailey serving as judge; George and Lexie adjust to their new, squalid apartment; Callie finds a friend in Erica.
The nurses' complaints about Mark Sloan's trysts result in a date-and-tell policy; Cristina is forced to face her feelings for Burke; Meredith and Derek admit a veteran into their clinical trial.
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