The nurses' complaints about Mark Sloan's trysts result in a date-and-tell policy; Cristina is forced to face her feelings for Burke; Meredith and Derek admit a veteran into their clinical trial.
Addison returns to Seattle Grace to perform an operation and is surprised by all the changes in the lives of her former co-workers; Derek and Meredith's clinical trial has its first patient; Rebecca returns with shocking news for Alex.
Dr. Wyatt refuses to let Meredith give up on therapy; Alex learns the truth about Rebecca's medical condition; the chief insists Erica perform a risky surgery on his mentor.
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