After learning that Arlo is a virgin, Maddie pays her understudy, Brittany, to seduce him. But things go awry when, after the act, Arlo says he is in love with Brittany and makes plans to spend the rest of his life with her.
After firing a stagehand, Maddie consoles him with a one night stand; Frank asks Arlo's crush to be his tango partner.
Maddie encounters the real Madison Banks (Kristen Johnston), a waitress whose audition (and name) she stole over 25 years ago.The real Maddie suggests that they exchange lives for an evening.
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